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The follow up question would be what us and isn't pain.
If a bacterium swimming in one direction encounters a toxin and changes direction to avoid dying, did it experience pain? If a tobacco plant reacts to attacks from insects by producing more nicotine and alerting its neighbours to do the same through signals sent through both rhizomes and airborne pheromones, does it experience pain? What about a worker ant, whose behaviour can be perfectly simulated by an algorithm simple enough that you can simulate hundreds of ants interacting?
Personally, I'd say none of these organisms are capable of feeling pain. Or if they are with the help of some definitions of what constitutes pain, it's just a signal like an automated assembly machine getting a signal from its sensors that a human entered its work space and it needs to slow down its robot arm to snails pace. So still incapable of suffering.
Also, if you set the threshold for what constitutes the ability for suffering too low, you quickly collide with the ethics of even early term abortion.